This is a very intriguing and useful thread. I moved my 5570 to the second slot and the correct power socket. The PCI fan seems pretty unpredictable - at the moment with just an audio sequencer running it's showing 1800. But if run the graphics stress thing it drops to 800. I wonder if there's a way to automatically trigger the Unreal Heaven thing at bootup...
Do you have an Apple OEM 5770? Are you using a Mac Pro 5,1 or 4,1 (or other)?
I would not have expected your PCI fan speeds to fluctuate like that in Slot 2. But, if you are using a 'supported' card, then perhaps Apple insist you have it in Slot 1.
I use an unsupported card, so I see odd fan behavior in Slot 1, but not in Slot 2.
Another Forum Member (Actionable Mango, I think) mentioned in another thread that this is known phenomenom. Unsupported cards will produce odd fan behavors in SLot 1, but you can get around that with certain 'tricks'. One trick is using Slot 2. The other is stressing the GPU with a benchmark so the PCI fan throttles back down to a steady 800 rpms.
The Stress-test trick does not survive a reboot, however, and you PCI fan will start fluctuating again.
In the end, I just returned the GTX 650 to Slot 2 where fanspeed is predictable and steady. It blocks access to SLot 3, but I've got no other cards in there.
I forgot to mention that Mango stated that some users of Mac Pro 4,1 have reported the fanspeed fluctuations with an Apple 5770 (which is not 'officially' supported on that model Mac Pro).