That's a complex question, and the best answer is probably "it depends."
If you have less than about 110GB of stuff, you'll never use your hard drive and everything will stay on the SSD. In that case, your experience will be that Fusion Drive is SSD-fast, all of the time.
If you have more than about 110GB of stuff, your most frequently-used data will end up being tiered to the SSD. If you've got a set of files you access regularly, those files will likely be promoted to SSD and stuff you access less-frequently will be demoted to HDD, so if you're going through a regular workflow with mostly the same applications and the same data files, they should end up on the SSD.
It's impossible to predict how "fast" Fusion Drive will be after the SSD fills up and it has to begin promoting and demoting things. Indications are that whatever algorithm Core Storage uses to track frequently accessed chunks is pretty sensitive to that frequency and that it's pretty aggressive in keeping hot data on the SSD. Also, anything new you add should land on the SSD first, so if you're working with audio and you add a bunch of new audio files from external media to your Fusion Drive, those new files will wind up on the SSD first (and something cold will be demoted from SSD to HDD to make room for them, because FD strives to keep a minimum of 4GB free on the SSD).
If, however, you really have >110GB of data and you need constant SSD-speed access to *all* of it, *all the time*, you'll probably notice quite a few hiccups as FD shuffles data around, with the hiccups occurring more frequently the more data you have. Someone with a monster-big Aperture library might run into this, for example (though I'd suggest to that person that they consolidate their originals onto an external drive or a network share to keep the library's size down).
Your answer to this question will be different from someone else's answer, and there's just no way to know until you have an idea of how much stuff you have and what your access patterns are going to be like. There's certainly no useful way to quantify it, like, "Fusion Drive will be X% as fast as having a giant SSD," because the subjective "speed" of FD is totally dependent on what files are on it, how full the SSD is, and what you're accessing when.