They are the sensors that will monitor heartbeat and other functions
Movement via accelerometer and possibly gyroscope.
Each sensor window contains one set of LEDs and photodiodes; a pair of sensors use green visible light LEDs with matching photodiodes, the other pair uses infra-red LEDs.Two are IR LEDs, and the other two are the actual optical sensors.
What other functions?
Accelerometers and gyroscopes are teeny-tiny components mounted on the PCB. What do the LEDs and light sensors have to do with this?