8 DIMM slots vs 4 DIMMs... and think in PAIRS.
You are paying $500 for 2 extra memory upgrades, or not having to toss out the memory you have in the machine when you want to upgrade it again after you used your first 3rd party pair.
This is most evident in the mini with 1 DIMM slot... 2 DIMMs would be better for mini users and 4 would be happy days.
The PM is quite similar basically being 2 pair or 4 pairs, though a PM with 1 pair would drive you nuts.
Larger HD, 2 extra memory upgrades, PCI-X (which you may not use), and a slight MHz bump.
RAM may not be important to you now, but in a couple years -- maxing out at 16GB may be worthwile.