Application sandboxing and mandatory access controls (MAC) in OS X are the same thing. More specifically, applications are sandboxed in OS X via MAC. Mac OS X uses the TrustedBSD MAC framework, which is a derivative of MAC from SE-Linux. This system is mandatory because it does not rely on inherited permissions. Both mandatorily exposed services (mDNSresponder, netbios...) and many client-side apps (
Safari, Preview, TextEdit…) are sandboxed in Lion.
Windows does not have MAC. The system that provides sandboxing in Windows, called mandatory integrity controls (MIC), does not function like MAC because it is not actually mandatory. MIC functions based on inherited permissions so it is essentially an extension of DAC (see #1). If UAC is set with less restrictions or disabled in Windows, then MIC has less restrictions or is disabled.
http://www.exploit-db.com/download_pdf/16031 -> article about Mac sandbox.
http://msdn.microsoft.com/en-us/library/bb648648(v=VS.85).aspx -> MS documentation about MIC.
https://media.blackhat.com/bh-eu-11/Tom_Keetch/BlackHat_EU_2011_Keetch_Sandboxes-Slides.pdf -> researchers have found the MIC in IE is not a security boundary.