HiI've never tried an iMac with an SSD before, but I commonly use Macbook Pro's with SSD's. For example, my friend has a 15" retina Macbook Pro (Iris Pro model from 2014)and I really don't tell that much of a difference.
Without a doubt, bouts times are insanely better (another price who just bought a 15" Macbook, I was hoping her set it up and we went to restart it, looked away for a second and it was done ) but when it came to normal tasks such as launching apps, it was not really as big a gap. The Disk Speed Test for example, take the exact same time to open on my iMac with a HDD (not cached in RAM, was after a restart) as a 15" retina Macbook Pro. Larger Pro apps are more noticeable, say a second or two, but this is for opening them the fist time after a reboot. Once an app is cached in RAM (opened before), there was absolutely 0 difference between an SSD Macbook Pro 15" (with a slightly faster CPU and slower GPU but with double the RAM) and my iMac.
I feel that the main selling point of an SSD is its silent running, and its lower failure rates with knocks, as the speed different is only really noticeable when booting up (with Mac OS I only ever restart once in a blue moon, and with my Windows partition for gaming, I just hibernate it so that I can boot into Windows in around 7 seconds).
There is only once main place (apart from booting) that it was noticeable though, and that is when doing multiple disk intensive tasks at the same time (for example, copying a large file as well as trying to open a large app, as well as rendering, will shown the difference much more, but I really don't think that is worth several hundred pounds).
How is a 1TB HDD upgrade from Apple?
To be honest, I don't notice it being any quieter. I'm sure it is quieter, but the fans are usually all I can hear on my laptop (non-retina - think newer ones are even quieter). I don't really restart either - my uptime at the moment is 10 days.
It is most noticeable on boot, but I'd disagree that that's the main place you feel the speed. Day to day, you would notice it. We have a shared iMac at home which has a 5400rpm HDD which used to be roughly the same speed as my MBP. Now, you compare the two and the difference is night and day. I find it hard to use now!
SSDs probably aren't a necessity, but I wouldn't dismiss them without living with one on your main machine for a good week or two. Going back to a spinning drive isn't a pleasurable experience! I was ready to return my SSD as I only bought it as an experiment and it was 512GB compared to my old 1TB drive, but I liked it so much I kept it. Should tide me over until Skylake!