Even on the native drive bays which are SATA II an SSD will make a big improvement. All you need is an adapter to fit SSD to the drive sled.
I would advise to not buy a 2008, you should be able to find a 2009 for $495 and up.
You can add SATA III PCIe cards designed for SSDs. Or there are PCIe adapters, not controllers, that let you install even faster (~2x+ what SATA III offers, faster than a pair of SSDs in RAID) using an M.2 blade SSD.
So for $75 + sled adapter you can have 128GB for the system and enjoy SSD.
For $500+ you can get an Apple or Samsung blade. And for $300 you could get a pair of 250GB SSDs and controller.
So at least 3 or 4 different options.