There's no such thing, as far as I know. There is, however, an ” that is HTML code for right double quotes. Perhaps you, or the one you are "quoting", has mistaken the "&" for an "E" (for lack of better context)?
The key used to initiate the grave accent is the same key used for the ~, at least on a US keyboard. That's the key immediately to the left of the number 1 - usually directly above the tab key.
I think one issue with THAT key, is that it might appear in other locations.
You can sort out where it will be on other keyboards with the Keyboard Viewer app, which is available through the Keyboard pref pane, at least in Yosemite.